The Anthropic Messages passthrough and the token-count forwarding both build their upstream request through `get_anthropic_request_target`, which sends the connection key as `Authorization: Bearer <key>`. Anthropic's OpenAI-compatible `/chat/completions` endpoint accepts that, which is why the model works in the chat UI, but the native `/v1/messages` and `/v1/messages/count_tokens` endpoints do not: they require the key in `x-api-key` and reject a bearer token with 401 `Invalid bearer token` (and `jwt auth is not yet supported on count_tokens`). They also require an `anthropic-version` header, which was never sent.
For `api.anthropic.com` connections, send `anthropic-version` and move the key into `x-api-key`, dropping the bearer header. Connections using session, OAuth or Entra ID auth keep their token untouched, LiteLLM passthrough connections are unaffected, and admin-configured custom headers still win over both defaults.
Fixes#27695
f8fe0cde9a